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Termites are eusocial insects that are categorized in the taxonomic rank of infraorder Isoptera, or as epifamily Termitoidae within the cockroach order Blattodea. Termites were once classified in another order from cockroaches, but recent phylogenetic studies indicate that they evolved from close ancestors of cockroaches during the Jurassic or Triassic.

Approximately 3,106 species are currently described, using a couple hundred more left to be clarified. Although these insects are often called"white ants", they are not ants. .

Like ants and some bees and wasps in the separate order Hymenoptera, termites divide labour among castes consisting of sterile male and female"workers" and"soldiers". All colonies have fertile men called"kings" and one or more fertile females called"queens". Termites mostly feed on dead plant material and cellulose, generally in the form of wood, leaf litter, dirt, or animal dung.

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Termites are among the most successful groups of insects on Earth, colonising most landmasses except Antarctica. Their colonies range in size from a couple hundred individuals to enormous societies using several million individuals. Termite queens have the longest lifespan of any insect in the world, with a few queens reportedly living up to 30 to 50 years.

Colonies are described as superorganisms because the termites form a part of a self-regulating entity: the colony itself. .

Termites are a delicacy in the diet of some human cultures and are employed in many traditional medicines. A couple hundred species are economically significant as insects which can cause serious damage to buildings, plants, or plantation forests. Some species, like the West Indian drywood termite (Cryptotermes brevis), are considered as invasive species. .

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The infraorder name Isoptera comes from the Greek words iso (equivalent ) and ptera (winged), which describes the nearly equal size of their fore and hind wings.2"Termite" derives from the Latin and Late Latin term termes ("woodworm, white ant"), altered by the influence of Latin terere ("to rub, wear, erode") from the earlier word tarmes.

The external appearance of this giant northern termite Mastotermes darwiniensis is indicative of the close relationship between termites and cockroaches.

Termites were formerly placed in the order Isoptera. As early as 1934 suggestions were made they were closely linked to wood-eating cockroaches (genus Cryptocercus, the woodroach) dependent on the similarity of their symbiotic gut flagellates.6 In the 1960s additional evidence supporting that hypothesis emerged when F. A. McKittrick noted similar morphological traits between some termites and Cryptocercus nymphs.7 In 2008 DNA analysis from 16S rRNA sequences8 supported the position of termites being nested within the evolutionary tree containing the sequence Blattodea, which included that the cockroaches.910 The cockroach genus Cryptocercus shares the strongest phylogenetical similarity with termites and is considered to be a sister-group to termites.1112 Termites and Cryptocercus share similar morphological and societal features: for instance, most cockroaches do not display social characteristics, but Cryptocercus takes care of its young and exhibits other societal behaviour such as trophallaxis and allogrooming.13 Termites are regarded as the descendants of the genus Cryptocercus.914 Some researchers have suggested that a more conservative measure of retaining the termites as the Termitoidae, an epifamily within the cockroach order, which preserves the classification of termites in family level and below.15 Termites have long been approved to be closely associated with cockroaches and mantids, and they are categorized in precisely the exact same superorder (Dictyoptera).1617.

The oldest unambiguous termite fossils date to the early Cretaceous, but given the diversity of Cretaceous termites and ancient fossil records showing mutualism between microorganisms and these insects, they probably originated earlier in the Jurassic or Triassic.181920 Further evidence of a Jurassic origin would be that the assumption that the extinct Fruitafossor consumed termites, judging from browse around this site the morphological similarity to modern termite-eating mammals.21 The oldest termite nest detected is thought to be by the Upper Cretaceous in West Texas, in which the earliest known faecal pellets were also discovered.22 Claims that termites emerged previously have confronted controversy.

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Weesner indicated the Mastotermitidae termites may go back to the late Permian, 251 million years ago,23 and fossil wings that have a close resemblance to the wings of Mastotermes of their Mastotermitidae, the most primitive living termite, have been found in the Permian layers in Kansas.24 It is even possible that the very first termites emerged during the Carboniferous.25 The folded wings of this fossil wood roach Pycnoblattina, arranged in a convex pattern between segments 1a and 2a, resemble those seen in Mastotermes, the only living insect with exactly the identical pattern.24 Krishna et al., however, consider that all of the Paleozoic and Triassic insects tentatively categorized as termites are in fact unrelated to termites and needs to be excluded from the Isoptera.26 The primitive giant northern termite (Mastotermes darwiniensis) exhibits numerous cockroach-like attributes which are not shared with other termites, like laying its eggs in rafts and having anal lobes on the wings.27 Cryptocercidae and Isoptera are united in the clade Xylophagidae.28 Termites are sometimes known as"white ants" but the only resemblance to the ants is because of their sociality which is because of convergent evolution2930 with termites being the very first social insects to evolve a caste system more than 100 million years ago.31 Termite genomes are generally relatively large compared to that of other insects; the first fully sequenced termite genome, of Zootermopsis nevadensis, which was published in the journal Nature Communications, consists of roughly 500Mb,32 while two subsequently published genomes, Macrotermes natalensis and Cryptotermes secundus, are considerably larger at around 1.3Gb.3330.

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